155,127 species and infraspecific names are in the database, 21,785 images, 59,958 bibliographic items, 434,141 distributional records.

Codium tomentosum Stackhouse

Classification:
Empire Eukaryota
Kingdom Plantae
Subkingdom Viridiplantae
Infrakingdom Chlorophyta infrakingdom
Phylum Chlorophyta
Subphylum Chlorophytina
Class Ulvophyceae
Order Bryopsidales
Family Codiaceae
Genus Codium

Pictures

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Codium tomentosum Stackhouse
Galicia, Spain, 2015. 24 Dec 2016. Ignacio Bárbara. © Ignacio Bárbara (barbara@udc.es).

Codium tomentosum Stackhouse
Galicia, Spain, 2015. 24 Dec 2016. Ignacio Bárbara. © Ignacio Bárbara (barbara@udc.es).

Codium tomentosum Stackhouse
Galicia, Spain, 2015. 24 Dec 2016. Ignacio Bárbara. © Ignacio Bárbara (barbara@udc.es).

Codium tomentosum Stackhouse
Codium tomentosum. 27 Dec 2013. Ignacio Bárbara. © Ignacio Bárbara (barbara@udc.es).

Codium tomentosum Stackhouse
upper subtidal, Ría de Camariñas, Galicia, Spain 2012. Ignacio Bárbara. © Ignacio Bárbara (barbara@udc.es).

Codium tomentosum Stackhouse
upper subtidal, Ría de Camariñas, Galicia, Spain 2012. Ignacio Bárbara. © Ignacio Bárbara (barbara@udc.es).

Codium tomentosum Stackhouse
upper subtidal, Islas Cíes, Galicia, Spain 2012. Ignacio Bárbara. © Ignacio Bárbara (barbara@udc.es).

Codium tomentosum Stackhouse
Collected near Roscoff (F), December 1984. 29 Dec 2011. Elmar Wosnitza. © Christine Wosnitza.

Codium tomentosum Stackhouse
Codium tomentosum, Spanish Point, Co. Clare, Ireland. 28 Aug 2010. M.D. Guiry. © Michael Guiry (mike.guiry@nuigalway.ie).

Codium tomentosum Stackhouse
Codium tomentosum, Spanish Point, Co. Clare, Ireland. 28 Aug 2010. M.D. Guiry. © Michael Guiry (mike.guiry@nuigalway.ie).

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Pocockiella variegata (J.V.Lamouroux) Papenfuss

Codium tomentosum Stackhouse Codium tomentosum, Spanish Point, Co. Clare, Ireland
© Michael Guiry (mike.guiry@nuigalway.ie)

Publication details
Codium tomentosum Stackhouse 1797: xxiv

Published in: Stackhouse, J. (1797). Nereis britannica; continens species omnes fucorum in insulis britannicis crescentium: descriptione latine et anglico, necnon iconibus ad vivum depictis... Fasc. 2. pp. ix-xxiv, 31-70, pls IX-XIII. Bathoniae [Bath] & Londini [London]: S. Hazard; J. White.
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Type species
This is the type species (holotype) of the genus Codium.

Status of name
This name is of an entity that is currently accepted taxonomically.

Origin of species name
Adjective (Latin), tomentose, i.e., thickly and evenly covered with short more or less adpressed curled or curved matted hairs (Stearn 1973).

Heterotypic Synonym(s)
Spongia dichotoma Hudson 1762
Fucus tomentosus Hudson 1778
Fucus tomentosus var. marginifer Turner 1811
Codium dichotomum var. ß marginiferum S.F.Gray 1821

Taxonomic notes
According to Silva et al. (1996) the name Codium dichotomum has been misapplied to this species by Børgesen (1949a: 22-23) and Brown & Jarman (1978: 1256).

Distributional notes
Although frequently reported for the Mediterranean Sea, the presence of this species in the area needs reassessment. Records of this species for Corsica, Sicily, Adriatic, Greece, Turkey, Libya and Tunisia probably refer to Codium vermilara (Gallardo et al., 1993). Records from the west coast of Africa should also be treated with suspicion (Lawson & Price, 1969; Lawson & John 1987 ).

Restricted to North Africa and Europe? (Silva et al., 1996; Verbruggen & Costa, 2015). - (26 Jun 2015) - Wendy Guiry

General environment
This is a marine species.

Common names

(as Codium tomentosum Stackhouse)
Indonesian: Soesoe lopek (Chapman & Chapman 1980).

Portuguese: Chorão, Chorão do mar, Pingarelhos (Oliveira, 1990).

Type information
Neotype locality: Acton Castle, Cornwall, England (Silva 1955: 571). Neotype: LINN (Silva 1955: 571). Notes: "Type. 'On the Devonshire and Cornwall Coasts: on the Long Rock between Marazion and Pensance1 plentiful. Hon. Mr. Wenman' (specimen not located). In the absence of the holotype, I have selected as neotype a specimen in the Linnean Society collected at Acton Castle, presumably by Stackhouse."

Detailed distribution with sources
(as Fucus tomentosus Hudson)
Africa: Ghana (Price, John & Lawson 1978).

(as Codium tomentosum Stackhouse)
Europe: Adriatic Sea (Giaccone 1978, Battelli & Vukovic 1995), Atlantic France (Burel, Le Duff & Ar Gall 2019), Balearic Islands (Rodríguez y Femenías 1889, Bellón 1921, Navarro & Bellón 1945), Britain (Newton 1931, Silva 1955, Burrows 1991, Hardy & Guiry 2003, Brodie et al. 2007, Brodie et al. 2016), Channel Islands (Lyle 1920, Dixon 1961, Trowbridge, Farnham & White 2004, Trowbridge & Farnham 2004), Corsica (Boudouresque & Perret 1977), France (Coppejans & Kling 1995, Dizerbo & Herpe 2007, Anon. 2017), France (Breizh) (Loiseaux-de Goër & Noailles 2008), Ireland (Adams 1907, Cotton 1912, Cotton 1913, Silva 1955, Parkes 1975, Guiry 1978, De Valéra et al. 1979, Burrows 1991, Morton 1994, Guiry 2012), Netherlands (Stegenga & Mol 1983, Stegenga et al. 1997), Portugal (Ardré 1970, Araujo et al. 2005, Araújo et al. 2009, Lopes et al. 2011), Spain (Lázaro Ibiza 1889, Sauvageau 1897, Bellón 1921, Hamel 1928, Miranda 1931, Bellón 1940, Bellón 1942, Ardré 1957, Fischer-Piette, C. & Seoane-Camba, J. (1962), Seoane-Camba 1965, Pérez-Cirera & Maldonado 1982, Conde Poyales & Seoane Camba 1982, Gallardo & Pérez-Cirera 1982, Fernández & Niell 1982, Anadón 1983, Gallardo et al. 1985, Anadón & Fernández 1986, Soto & Conde 1989, Pérez-Cirera 1989, Perez-Cirera, Cremades, Bárbara & Lopez 1991, Granja, Cremades & Barbara 1992, Gallardo et al. 1993, Flores-Moya et al. 1995, Bárbara, Cremades & Pérez-Cirera 1995, Bárbara & Cremades 1996, Veiga, Cremades & Bárbara 1998, Peña & Bárbara 2002, Valenzuela Miranda 2002, Sánchez, Fernández & Rico 2003, Gorostiaga et al., 2004, Bárbara et al. 2005, Peña & Bárbara 2006, Martínez-Gil et al. 2007, Peña & Bárbara 2008, Cires Rodriguez & Cuesta Moliner 2010, Hernández et al. 2010), Spain (incl. Canary Is.) (Gallardo et al. 2016), Turkey (Europe) (Güven & Öztig 1971, Güner, Aysel, Sukatar & Öztürk 1985, Cirik, Zeybeck, Aysel & Cirik 1990, Zeybek, Güner & Aysel 1993, Taskin et al. 2008 ).

Atlantic Islands: Azores (Tittley & Neto 1994, Parente), Canary Islands (Børgesen 1925, Gil-Rodríguez & Afonso-Carrillo 1980, Viera-Rodriguez et al. 1987, Gil-Rodriguez, Afonso-Carrillo & Wildpret de la Torre 1987, Haroun et al. 2002, Gil-Rodríguez et al. 2003, John et al. 2004, Anon. 2011, Afonso-Carrillo 2014), Cape Verde Islands (John et al. 2004, Prud'homme van Reine, Haroun & Kostermans 2005), Madeira (Levring 1974, Neto, Cravo & Haroun 2001, John et al. 2004), Salvage Islands (Parente et al. 2000, John et al. 2004).

North America: Florida (FL) (Taylor 1928), Gulf of California (Setchell & Gardner 1924).

South America: Argentina (Croce et al. 2015), Brazil (Taylor 1930), Chile (Ramírez & Santelices 1991), Peru (Ramírez & Santelices 1991), Temperate South America (Ramírez & Santelices 1991).

Africa: Angola (John et al. 2004), Congo (John et al. 2004), Egypt (Papenfuss 1968 , Aleem 1993), Ethiopia (Papenfuss 1968 ), Gabon (John et al. 2004), Kenya (Silva, Basson & Moe 1996, Bolton, Oyieke & Gwanda 2007), Madagascar (Silva, Basson & Moe 1996), Mauritania (John et al. 2004), Mauritius (Silva, Basson & Moe 1996), Morocco (Gallardo et al. 1993, Benhissoune, Boudouresque & Verlaque 2001, Benhissoune, Boudouresque & Verlaque 2001, Moussa et al. 2018), São Tomé & Príncipe (John et al. 2004), Senegal (John et al. 2004), Somalia (Silva, Basson & Moe 1996), South Africa (Silva, Basson & Moe 1996), Spanish North Africa (González García, Paredes Ruiz & Enrique Mirón 2017), Sudan (Papenfuss 1968 , Osman & Mohammed 2016), Tanzania (incl. Zanzibar) (Silva, Basson & Moe 1996), Tunisia (Ben Maiz, Boudouresque & Quahchi 1987).

Indian Ocean Islands: Amirante Islands (Silva, Basson & Moe 1996), Andaman Islands (Silva, Basson & Moe 1996), Laccadive Islands (Silva, Basson & Moe 1996), Réunion (Silva, Basson & Moe 1996), Rodrigues Island (Silva, Basson & Moe 1996, De Clerck et al. 2004), Seychelles (Silva, Basson & Moe 1996).

South-west Asia: India (Silva, Basson & Moe 1996, Gupta 2012, Rao & Gupta 2015), Israel (Einav 2007, Einav & Israel 2008), Jordan (Papenfuss 1968 ), Lebanon (Lakkis 2013), Sri Lanka (Silva, Basson & Moe 1996), Turkey (Asia) (Taskin et al. 2008 ), Yemen (Silva, Basson & Moe 1996).

Asia: Japan (Holmes 1896, Okamura 1936), Korea (Lee & Kang 2001, Bae 2010), South China Sea (Phang et al. 2016), Taiwan (Shao 2003-2014, Anon. 2012).

South-east Asia: Indonesia (Silva, Basson & Moe 1996, Atmadja & Prud'homme van Reine 2014), Philippines (Ang, Sin Man Leung & Mei Mei Choi 2014), Singapore (Silva, Basson & Moe 1996, Pham et al. 2011), Vietnam (Pham-Hoàng 1969, Tien 2007, Nguyen et al. 2013).

Australia and New Zealand: Queensland (Lewis 1987 ).

Pacific Islands: Central Polynesia (Tsuda & Walsh 2013), Samoan Archipelago (Skelton & South 1999).

Antarctic and the subantarctic islands: Antarctica/Subantarctic Islands (Papenfuss 1964).

(as Fucus tomentosus var. marginifer Turner)
Asia: Taiwan (Shao 2003-2014).

Key references
Cormaci, M., Furnari, G., & Alongi, G. (2014). Flora marina bentonica del Mediterraneo: Chlorophyta. Bollettino dell'Accademia Gioenia di Scienze Naturali di Catania 47: 11-436.

Kim, G.H. & Klochkova, T.A. (2010). Bryopsidales. In: Algal flora of Korea. Volume 1, Number 1. Chlorophyta: Ulvophyceae: Ulotrichales, Ulvales, Cladophorales, Bryopsidales. Marine green algae. (Bae, E.H., Kim, H.-S., Kwon, C.-J., Hwang, I.-K., Kim, G.H. & Klochkova, T.A. Eds), pp. 157-209. Incheon: National Institute of Biological Resources.

Loiseaux-de Goër, S. & Noailles, M.-C. (2008). Algues de Roscoff. pp. [1]-215, col. figs. Roscoff: Editions de la Station Biologique de Roscoff.

Newton, L.M. (1953). Marine algae. Scientific Reports of the John Murray Expedition 1933-34 9(5): 395-420, pls I-IV.

Nizamuddin, M. (1991). The green marine algae of Libya. pp. 1-230. Bern: El-Faateh University, Department of Botany.

Zemke-White, W.L. & Ohno, M. (1999). World seaweed utilisation: an end-of-century summary. Journal of Applied Phycology 11: 369-376.

Created: 06 April 1996 by M.D. Guiry

Verified by: 03 February 2017 by M.D. Guiry

Accesses: This record has been accessed by users 23479 times since it was created.

Verification of data
Users are responsible for verifying the accuracy of information before use, as noted on the website Content page.

References
(Please note: only references with the binomials in the title are included. The information is from the Literature database.)

Cires Rodriguez, E. & Rico Ordás, J.M. (2007). Distribución, crecimiento y fenología del alga invasora Codium fragile y su congénere nativa Codium tomentosum. In: (Real Instituto de Estudios Asturianos, eds.) I Congreso de Estudios Asturianos (2006) Tomo VI. Comisión de Ciencias de la Naturaleza y Tecnología, Oviedo, pp. 145-161. ( Eds)
Hassoun, M., Salhi, G., Boukssir, H., Moussa, H., Riadi, H. & Kazzaz, M. (2015). Codium tomentosum var. mucronatum et son epiphyte Aglaothamnion pseudobyssoides, deux nouvelles espèces d'algues benthiques pour la phycoflore du Maroc. Acta Botanica Malacinata 39: 37-44.
Tobler, G. (1911). Zur organisation des thallus von Codium tomentosum. Flora 103: 78-87.
Trowbridge, C.D. (2001). Coexistence of introduced and native congeneric algae: Codium fragile and C. tomentosum on Irish rocky intertidal shores. Journal of the Marine Biological Association of the United Kingdom 81: 931-937.
Trowbridge, C.D., Farnham, W.F. & White, L.F. (2004). Thriving populations of the native macroalga Codium tomentosum on Guernsey rocky shores. Journal of the Marine Biological Association of the United Kingdom 84: 873-877.
Went, F.A.F.C. (1889). Les modes de reproduction du Codium tomentosum. Nedserl. Kruidk. Archf. Ser. 2, 5: 440-444.
Williams, M.M. (1925). Contributions to the cytology and phylogeny of the siphonaceous algae. I. The cytology of the gametangia of Codium tomentosum (Stack.). Proceedings of the Linnean Society of New South Wales 50: 98-111.

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Cite this record as:
M.D. Guiry in Guiry, M.D. & Guiry, G.M. 2019. AlgaeBase. World-wide electronic publication, National University of Ireland, Galway. http://www.algaebase.org; searched on 25 April 2019.

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