156,011 species and infraspecific names are in the database, 21,990 images, 60,295 bibliographic items, 442,406 distributional records.

Melobesia Heydrich, 1897, nom. illeg.

Empire Eukaryota
Kingdom Plantae
Subkingdom Biliphyta
Phylum Rhodophyta
Subphylum Eurhodophytina
Class Florideophyceae
Subclass Corallinophycidae
Order Hapalidiales
Family Hapalidiaceae
Subfamily Melobesioideae

Holotype species: Melobesia pustulata J.V.Lamouroux

Currently accepted name for the type species: Lithophyllum pustulatum (J.V.Lamouroux) Foslie

Original publication:Heydrich, F. (1897). Melobesiae. Berichte der deutsche botanischen Gesellschaft 15: 403-420, pl. XVIII [18].
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Type designated in Foslie, M. (1898). Systematical survey of the Lithothamnia. Det Kongelige Norske Videnskabers Selskabs Skrifter 1898(2): 1-7.
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Precise date of publication7 September 1897 (see Woelkerling & Lamy 1998: 680 for details). The requirements for valid publication are specified in the ICBN (International Code of Botanical Nomenclature)

Taxonomic status: currently regarded as a synonym of Lithophyllum.

Most recent taxonomic treatment adopted: Brook, A.J. & Johnson, L.R. (2002). Order Zygnematales. In: The Freshwater Algal Flora of the British Isles. An identification guide to freshwater and terrestrial algae. (John, D.M., Whitton, B.A. & Brook, A.J. Eds), pp. 479-593. Cambridge: Cambridge University Press.

Nomenclatural notes
The ICBN rules cited below are those adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005 (McNeill et al. 2006).

In accordance with ICBN Art 13.3, the name Melobesia Heydrich is treated as pertaining to a non-fossil taxon because its type is based on a non-fossil specimen. No species based on fossil types have been assigned to the genus.

The generic name Melobesia Heydrich is typified (ICBN Art 10.1) by the type of M. pustulata, the designated lectotype species of the genus and one of ten species originally (Heydrich 1897) included in the genus. Although validly published, Melobesia Heydrich (1897) is an illegitimate later homonym (ICBN Art. 53) of Melobesia Lamouroux (1812). Consequently, Melobesia Heydrich (1897), which is not conserved, must be rejected in accordance with ICBN Art. 45.3. - (22 Jul 2009) -

Description: No description is provided as this name is currently treated as a synonym.

Information kindly contributed by but may now be outdated.

Comments: Heydrich (1897: 408) excluded the type of Melobesia Lamouroux (M. membranacea) from his genus by placing it in his hew genus Epilithon (see separate AlgaeBase entry), thereby establishing a new genus called Melobesia which must be solely attributed to him. Heydrich characterized his genus as having uniporate tetrasporangial conceptacles rather than multiporate tetrasporangial conceptacles as occur in Melobesia membranacea Lamouroux. Heydrich did not designate a type species, but Foslie (1898a: 6), who briefly accepted Heydrich’s concept of Melobesia, designated M. pustulata Lamouroux as type. The type of Melobesia pustulata is also the type species of the older generic name Titanoderma Nägeli (1858: 532), which currently is either recognized as a distinct genus or treated as a heterotypic synonym of Lithophyllum Philippi (1837). Additional information about Melobesia Heydrich is provided by Mason (1953: 320) and Woelkerling (1988: 113).

Heydrich’s concept of Melobesia, or that of Foslie (1898b: 10), who established a second later homonym (Melobesia Foslie; see separate AlgaeBase entry), persisted, at least in part, in the literature for many years (e.g. see Levring 1974; Etcheverry 1986) even though Mason (1953) had clearly summarized the ‘illegal’ (=illegitimate) nature of the Heydrich and Foslie names.

Although the generic name Melobesia Heydrich is illegitimate, all ten species names that Heydrich (1897: 408-409) included in the original presentation of Melobesia Heydrich are legitimate (see Art. 55 of the International Code of Botanical Nomenclature.

Numbers of names and species: There is only one species or infraspecific name in the database at present.

Names: ('C' indicates a name that is accepted taxonomically; 'S' a homotypic or heterotypic synonym; 'U' indicates a name of uncertain taxonomic status, but which has been subjected to some verification nomenclaturally; 'P' indicates a preliminary AlgaeBase entry that has not been subjected to any kind of verification. For more information on a species click on it to activate a link to the Species database):

Click here to also show infraspecific names in the list below.

Etcheverry, D.H. (1986). Algas marinas bentónicas de Chile. pp. [1]-379, illus.. Montevideo, Uruguay: UNESCO (Regional office of Science and Technology for Latin America and the Caribbean).

Foslie, M. (1898). Systematical survey of the Lithothamnia. Det Kongelige Norske Videnskabers Selskabs Skrifter 1898(2): 1-7.

Foslie, M. (1898). List of species of the Lithothamnia. Kongelige Norske Videnskabers Selskabs Skrifter 1898(3): 1-11.

Foslie, M. (1904). Die Lithothamnien des Adriatischen Meeres und Marokkos. Wissenschaftliche Meeresuntersuchungen, Abt. Helgoland, N. F. 7(1): 1-40, pls 1-3.

Heydrich, F. (1897). Melobesiae. Berichte der deutsche botanischen Gesellschaft 15: 403-420, pl. XVIII [18].

Lamouroux, [J.V.F.] (1812). Extrait d'un mémoire sur la classification des Polypiers coralligènes non entièrement pierreux. Nouveaux Bulletin des Sciences, par la Société Philomathique de Paris 3: 181-188.

Lamouroux, J.V.F. (1816). Histoire des polypiers coralligènes flexibles, vulgairement nommés zoophytes. pp. [i]-lxxxiv, chart, [1]-560, [560, err], pls I-XIX, uncol. by author. Caen: De l'imprimerie de F. Poisson.

Levring, T. (1974). The marine algae of the Archipelago of Madeira. Boletim Museo Municipale Funchal 28: 5-111.

McNeill, J., Barrie, F.R., Burdet, H.M., Demouline, V., Hawksworth, D.L., Marhold, K., Nicolson, D.H., Prado, J., Silva, P.C., Skog, J.E., Wiersema, J.H. & Turland, N.J. (2006). International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botancial Congress Vienna, Austria, July 2005. pp. [i-iv], v-xviii + 1-568. Liechtenstein: A.R.G. Gantner Verlag.

Mason, L.R. (1953). The crustaceous coralline algae of the Pacific coast of the United States, Canada and Alaska. University of California Publications in Botany 26: 313-389, Plates 27-46.

Nägeli, C. (1858). Die Stärkekörner. In: Pflanzenphysiologisce Untersuchungen. (Nägeli, C. & Cramer, C. Eds) Vol. 2, pp. i-x, 1-623. Zürich:

Philippi, R.A. (1837). Beweis, dass die Nulliporen Pflanzen sind. Archiv für Naturgeschichte 3: 387-393, figs 2-6, pl. IX.

Woelkerling, W.J. (1988). The Coralline Red Algae: an analysis of the genera and subfamilies of nongeniculate Corallinaceae. pp. i-xi, 1-268, 259 figs, tables numbered by chapter. London & Oxford: British Museum (Natural History) & Oxford University Press.

Woelkerling, W.J. & Lamy, D. (1998). Non-geniculate coralline red algae and the Paris Museum: Systematics and scientific history. pp. i-vii, 1-767. Paris: Publications Scientifiques du Muséum/ADAC.

Verification of data
Users are responsible for verifying the accuracy of information before use, as noted on the website Content page.

Some of the descriptions included in AlgaeBase were originally from the unpublished Encyclopedia of Algal Genera, organised in the 1990s by Dr Bruce Parker on behalf of the Phycological Society of America (PSA) and intended to be published in CD format. These AlgaeBase descriptions are now being continually updated, and each current contributor is identified above. The PSA and AlgaeBase warmly acknowledge the generosity of all past and present contributors and particularly the work of Dr Parker.

Descriptions of chrysophyte genera were subsequently published in J. Kristiansen & H.R. Preisig (eds.). 2001. Encyclopedia of Chrysophyte Genera. Bibliotheca Phycologica 110: 1-260.

Created: 22 July 2009 by William J. Woelkerling

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Guiry, M.D. & Guiry, G.M. 2019. AlgaeBase. World-wide electronic publication, National University of Ireland, Galway. http://www.algaebase.org; searched on 19 August 2019.

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